Graduation Date

Spring 2025

Document Type

Thesis

Program

Master of Science degree with a major in Biology

Committee Chair Name

Rachael Wade

Committee Chair Affiliation

Cal Poly Humboldt Faculty or Staff

Second Committee Member Name

Sean Craig

Second Committee Member Affiliation

Cal Poly Humboldt Faculty or Staff

Third Committee Member Name

Mark Carr

Third Committee Member Affiliation

Community Member or Outside Professional

Fourth Committee Member Name

Brent Hughes

Fourth Committee Member Affiliation

Community Member or Outside Professional

Keywords

Northern California, Mendocino, River mouth, Bull kelp, Purple urchins, Refugia, Nereocystis luetkeana, Marine heatwaves, Atmospheric rivers, Climate change, Kelp forest, Urchin barren

Subject Categories

Biology

Abstract

Populations of Nereocystis luetkeana (bull kelp) have experienced localized extinction events in northern California since 2014 due to multiple stressor events, including the area’s most severe marine heatwave (MHW) to-date. Previous surveys in the region observed that bull kelp recovered and persisted at river mouths following the 2014-2015 MHW, suggesting that river mouths may serve as refugia – the refugial river mouth hypothesis. My study aimed to assess this hypothesis and the long-term stability of these refugia by studying how freshwater influences communities at these sites and bull kelp recruitment. Field surveys of salinity fluctuation and kelp and sea urchin densities were conducted at paired river mouth and coastal reference sites. A laboratory salinity pulse experiment examined gametophyte and sporophyte development to simulate freshwater discharge events. River plume models suggested that river mouths experienced reductions in salinity in response to high flow events (minimum of 20 PSU), such as storm events, while reference sites did not experience such fluctuations. Survey results showed significantly higher abundances of urchins at reference sites than at the river sites, with bull kelp persisting at comparable abundances at all river sites and some reference sites. Kelp gametophytes survived repeated salinity pulses as low as 8 PSU, while healthy sporophytes were only observed at or above 16 PSU. Thus, river mouths are likely not limiting bull kelp recruitment, development and survival, but are reducing herbivore pressure. These results support the refugial river mouth hypothesis and that these sites will likely continue to harbor bull kelp and should be targeted for restoration efforts.

Citation Style

APA

Available for download on Tuesday, May 12, 2026

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